What Do Swifts Eat?

samples of swift chick droppings were collected at various sites across Northern Ireland between 2010 and 2022. My House, A Birch Hill Park, will be a permanent year on year study. Droppings in all cases were collected off the ground below nests when chicks were c3 weeks old and can poop out of nest sites. Droppings were dried and sent away for analaysis of insect contents

No further classification and eg mean the insect fragments couldnt be IDd to species

Swift site locations

A - collected at Birch Hill Park BT41 1DE

Country swifts living beside Lough Neagh – my house 1 mile / 1.6km from the shore of Lough Neagh.

Chironomids 25.7% - non biting midges e.g. Lough Neagh fly
Aphids 18.0% - greenfly, blackfly
Psyllids 11.6% - sap sucking insects
Lonchoptera 11.5% - small spear-winged flies
Coleoptera 11.1% - water beetles
Phoridae 0.8% - hump-backed flies (resembling fruit flies)
Sciaridae 0.7% - fungus gnats
Dolichopodidae 0.4% - long-legged flies
Muscids / Calliphorids 0.4% - house / stable / blow flies
Scathophagidae 0.3% - dung flies
Hemiptera 2.1% - bugs
Tipulidae 0.1% - craneflies

with traces of
Hymenoptera - small solitary wasps
Coccinellidae - 11 spot ladybird

Birch Hill Park BT41 1DE 2021

  • Chironomidae non-biting midges .66%
    Tipulidae craneflies 4.95%
    Diptera (S.O. Nematocera) No further classification 4.29%
    Scathophagidae dungflies 10.9%
    Calliphoridae blow flies 1.65%
    Muscidae house flies 26%
    Diptera (S.O. Cyclorrhapha) No further classification 11.90%
    Scarabaeidae chafers 12.89%
    Coccinellidae ladybirds 1.32%
    Coleoptera beetles No further classification 10.24%
    Aphidae aphids 20.49%
    Cercopidae froghoppers 5.95%
    Cicadellidae leafhoppers 1.98%
    Hemiptera true bugs No further classification 9.91%
    Ichneumonidae Ichneumons 6.4%
    Cynapidae gall wasps 5.61%
    Hymenoptera wasps
    No further classification 8.26%
  • Birch Hill Park BT41 1DE 2022
  • Chironomidae non-biting midges .13%
    Tipulidae craneflies 4.05%
    Nematocera eg knats and midges 1.12%
    Scathophagidae dungflies 8.91%
    Calliphoridae blow flies 1.35%
    Muscidae house flies 3.51%
    Cyclorrhapha eg circular seamed flies 9.72%
    Scarabaeidae chafers 10.54%
    Coccinellidae ladybirds 1.08%
    Coleoptera beetles No further classification 8.37%
    Aphidae aphids 16.75%
    Cercopidae froghoppers 4.86%
    Cicadellidae leafhoppers 1.62%
    Hemiptera (true bugs) No further classification 8.10%
    Ichneumonidae Ichneumons 5.67%
    Cynapidae gall wasps 4.59%
    Hymenoptera wasps
    No further classification 6.75%
  • Comments from the person doing the analysis
    The main insect groups in the sample were like those in the 2019 and 2021 samples from the
    same site although, as may be expected in a different summer, the relative proportions varied.
    Noticeable changes were the increased proportion of Nematocera in the pellets this year. The
    summer in the area having been less dry, the number of longhorn flies is likely to have been
    higher than in 2021; in fact the numbers of longhorn fragments seems to have returned to a
    level similar to that in 2019, i.e. to a more normal year. Cyclorrhaphan flies were also more
    numerous in the diet in 2022, probably for similar reasons. The numbers of beetles were
    similar to those in 2021; again, the main group was small chafers (Serica sp) which hatch in
    large numbers from plants and get carried upwards on air currents. Their numbers were
    broadly similar to those in 2021. It was noticeable that the numbers of hemipteran bugs were
    down this year, mainly due to the drop in the proportion of aphids. Conditions may have been
    less suitable for aphids, or it could be that there were fewer strong up currents taking them to
    the height at which swifts forage. The reduction in the proportion of wasps may also have
    been due to fewer strong up currents.
    Overall, the diet of swifts at this site is pretty constant from year to year, and the proportions
    of different groups was generally more similar this year to 2019 than to 2021. This is likely to
    be due to weather conditions rather than to major changes in the insect population.
  • Birch Hill Park BT41 1DE 2023
    This year while collecting droppings part of a bolus was found
    Bolus results
    Cynapidae (gall wasps) 19.7%
    Aphidae (aphids) 18.5%
    Chironomidae (non-biting midges) 18.5%
    Ichneumonidae (Ichneumons) 14.8%
    true bugs No further classification 12.3%
    Nematocera (midges, gnats, mosquitoes) No further classification 7.4%
    Cyclorrhapha (house fly types) No further classification 6%
    Cyclorrhapha Muscidae 2.5%
    Total number of identifiable items - 81

    Comments
    Clearly, on the day the bird collected this bolus, Nematocera were in abundance. This
    analysis should be regarded as a snapshot of what a bird found to eat on one day, rather than a
    detailed diet analysis. It further shows that swifts, like most animals, feed on what they can
    get at the time.

    Droppings results

    (true bugs) No further classification 20.2%
    Aphidae (aphids) 10.5%
    Cyclorrhapha (circular-seamed flies) No further classification 10.7%
    Scarabaeidae (chafers) 10.22%
    Hymenoptera (wasps) No further classification 9%
    Lonchopteridae (spear-winged flies) 6.73%
    Cynapidae (gall wasps) 6.73%
    Chironomidae (non-biting midges) 6%
    Miridae (capsid bugs) 4.45%
    Sciomyzidae (marsh flies) 3.74%
    Ichneumonidae (Ichneumons) 3.74%
    Scathophagidae (dung-flies) 3.49%
    Neuroptera (lacewings) Hemerobiidae (brown lacewings) 2%
    Coleoptera (beetles) No further classification .75%
    Lepidoptera (moths) No further classification .75%
    Culicidae (mosquitoes) .74%
    Neuroptera (lacewings) No further classification .2%

    Total number of identifiable fragments - 401

    Comments
    As in previous years, the main prey groups from this site were hemipteran bugs, cyclorraphan
    flies and wasps, and the relative proportions did not vary much between 2022 and 2023.
    Again as last year, nematoceran flies made up a smaller proportion of the diet – 6.7% this
    year. The proportion of beetles was down slightly – probably due to smaller rises of Serica
    chafers this summer. A similar drop in chafer numbers was noted at the other site examined
    this year – at CAFRE College. Wasp numbers were up on last year and are now more similar
    to those recorded in 2019 and 2021. Also as at CAFRE College, lacewing remains were
    recorded in the diet for the first time this year. Although the proportion was small, lacewings
    are not numerous insects, and their inclusion at all is an indication of larger numbers of
    lacewings in the area this year and/or strong upcurrents carrying them to greater heights.
    Moths were recorded in very small numbers. This has been noted in previous years, and the
    low proportion probably makes their inclusion of little importance to the swift population
    locally.
    As noted last year, the diet of the swifts at this site continues to be constant, taking into
    account small variations due to weather conditions.

    Birch Hill Park BT41 1DE 2024
    Bolus analysis
    Phoridae scuttle flies 28%
    Scathophagidae dung-flies 23%
    Chironomidae chironomids 12.4%
    Tipulidae craneflies 7.4%
    Cyclorrhapha circular-seamed flies No further classification 7.4%
    Dolichopodidae 4%
    Muscidae house flies 4%
    Culicidae mosquitos .23%
    Trichoceridae winter gnats .23%
    Number of items 121
    Comments
    It was notable that all the insects in the sample were flies and many of them dung flies, which
    feed on animal dung. While individuals were not identified to species, many of the dung flies
    were clearly the yellow dung fly (Scathophaga stercoraria) which feed in large numbers on
    cattle dung. Scuttle flies (Phoridae) have larvae which feed in decaying organic matter and
    are also common around cattle pastures. It is very likely, therefore, that the swift which
    produced this bolus was feeding over cattle pasture. Given the relatively large numbers of
    insects from a couple of families, this looks like opportunistic foraging on locally plentiful
    insects.
    This result differs from those of the faecal pellet analysis for the same period (see separate
    report), which found fewer flies in the diet and as many bugs, beetles and wasps as in
    previous summers. This difference emphasizes that the contents of a bolus will represent one
    foraging period only, as opposed to feeding over several days. The contents of the bolus are
    therefore likely to represent a relatively short foraging period and a more restricted location.
    Droppings analysis
    Scarabaeidae chafers 13.9%
    Hemiptera true bugs No further classification 9.4%
    Cyclorrhapha circular-seamed fly No further classification 9%
    Aphidae aphids 8.9%
    Scathophagidae dung-flies 8.2%
    Cynapidae gall wasps 8.2%
    wasps No further classification 7.3%
    Coleoptera beetles No further classification 5.6%
    Chironomidae non-biting midges 4.45%
    Miridae capsid bugs 3.6%
    Trichoptera (caddis flies) No further classification 3.6%
    Ichneumonidae Ichneumons 3.6%
    Pteromalidae Pteromid (parasitic) wasps 3.2%
    Lonchopteridae spear-winged fly 2.5%
    Muscidae house fly etc 2.5%
    Tipulidae craneflies .7%
    Diptera (S.O.Nematocera) No further classification .5%
    Delphacidae plant hopper 2%
    Phoridae scuttle bugs 1.5%
    Cicadellidae plant hoppers 1.3%

    Total number 584
    Comments
    Overall, diet composition this year was consistent with that in previous years at this site. As
    before, the main prey groups were hemipteran bugs, cyclorraphan flies and wasps. These
    three Orders, together with Coleoptera (most of which were small flying chafers (Serica spp)
    made up just over 90% of the diet. The relative proportions of the four groups were more
    even this year, with no group making up a huge proportion of the diet. As in past years, minor
    components of the diet were nematoceran flies and other orders, in this case caddis flies.
    While there were no real surprises in the results this year, four minor points could be
    considered:
    The proportion of beetles was higher than in 2023 and close to the value recorded in 2022
    (20%). This is in keeping with known behaviour of small, flying chafers which hatch in large
    “rises” periodically. This sample clearly coincided with a rise of Serica beetles, as in 2022.
    The proportion of the diet made up of cyclorraphan flies (many of them dung flies) was much
    the same as in previous years. However, as an aside, this year I noted that some of the faecal
    pellets in the study contained large numbers of fly remains (two of them contained very little
    else), while others contained none at all. This suggests that the flies were periodically available
    in large numbers but that at other times the birds did not find any. This could be due to the very
    changeable weather conditions prevailing in 2024. It also ties in with the results of the bolus
    analysis (see separate report), which found that the bolus contained only flies – clearly collected
    on a good “fly day”.
    Recent media reports have suggested that weather conditions nationally this summer may have
    been unfavourable for Hymenoptera as a group, since it appears there have been fewer social
    wasps around than in a more “normal” summer. This is clearly not the case here, since the
    proportion of Hymenoptera in the swifts’ diet is the highest yet recorded (22.4%). As before,
    most species eaten were solitary wasps with a parasitic phase. Their ecology is therefore
    different to that of social wasps.
    As in previous years, the largest component of the birds’ diet was hemipteran bugs, including
    aphids, which seem to have been as numerous as ever this year.
  • ______________________________________________________________________________________________________________________________
  • B - collected at the Crescent Arts Centre
    City swifts living in the Crescent Arts Centre, Belfast - 20km / 12 miles from Lough Neagh
    Thanks to Sue Swift for analysing insect fragments in the droppings
     
    Chironomidae non-biting midges 16.8%
    Hemiptera true bugs No further classification 12.0%
    Scathophagidae dung flies 11.8%
    Aphidae aphids 10.9%
    Ichneumonidae 7.6%
    Muscidae/Calliphoridae 7.4%
    Psyllidae jumping plant lice 7.3%
    Lonchopteridae  6.8%
    Platygasterodae 4.6%
    Scarabaeidae chafers 3.3%
    Sciomyzidae marsh flies 1.7%
    Tipulidae craneflies 1.7%
    Coleoptera beetles No further classification 1.6%
    Lepidoptera moths No further classification 0.9%
    Coccinellidae ladybirds 0.9%
    Cynapidae gall wasps 0.9%
    Delphacidae plant hoppers 0.9%
    Hymenoptera (wasps) No further classification 0.8%
    Phoridae 0.6%
    Diptera (S.O. Cyclorrhapha) No further classification 0.6%
    Hydroptilidae water beetles 0.6%
    Dytiscidae water beetles 0.3%
    ___________________________________________________________________________________________________________________

    C
    Droppings collected from a swift colony in Dervock in 2018
    Thanks to Sue Swift for analysing insect fragments in the droppings
     
    Hymenoptera (wasps) Ichneumonidae 14.26%
    Diptera (S.O. Cyclorrhapha) Muscidae (house flies) 11.93%
    Coleoptera (beetles) Scarabaeidae (chafers) 11.78%
    Hemiptera (true bugs) No further classification 10.54
    Diptera (S.O.Nematocera) No further classification 9.61%
    Coleoptera (beetles) No further classification 6.51%
    Diptera (S.O.Nematocera) Tipulidae (craneflies) 5.58%
    Hemiptera (true bugs) Aphidae (aphids) 4.80%
    Diptera (S.O. Cyclorrhapha) circular seamed flies No further classification 3.56%
    Neuroptera (lacewings) Hemerobiidae (brown lacewings) 3.10%
    Hemiptera (true bugs) Cicidellidae leaf hopper 2.63%
    Hymenoptera (wasps) Cynapidae (gall wasps) 2.94%
    Diptera (S.O.Nematocera) Chironomidae (non-biting midges) 2.63%
    Diptera (S.O. Cyclorrhapha Calliphoridae (blowflies) 2.01%
    Lepidoptera (moths) No further classification 1.39%
    Hemiptera (true bugs) Psyllidae (jumping plant lice) 1.39%
    Hymenoptera (wasps) No further classification 1.24%
    Diptera (S.O. Cyclorrhapha) Lonchopteridae spear-winged flies 1.24%
    Diptera (S.O. Cyclorrhapha) Sciomyzidae marsh flies .77%
    Diptera (S.O. Cyclorrhapha) Opomyzidae .77%
    Diptera (S.O. Cyclorrhapha) Syrphidae (hoverflies) .46%
    Diptera (S.O.Nematocera) Scatopsidae dung midges 3.46%
    ______________________________________________________________________________________________________________________________________________________________

    D droppings collected at St Naile's Catholic Church, Kinawley County Fermanagh
    collected June 24th 2019 4km to Lough Erne / 650m to Cladagh River
     
    Diptera (S.O. Cyclorrhapha) No further classification 121
    Hemiptera (true bugs) No further classification 100
    Diptera (S.O.Nematocera)  mosquitoes, crane flies, gnats, black flies 65
    Trichoptera (caddis flies) No further classification 48
    Scathophagidae (dung-flies) 44
    Muscidae (house flies) 40
    Tipulidae (craneflies) 37
    Chironomidae (non-biting midges) 36
    Hemiptera (true bugs) Aphidae (aphids) 34
    Diptera (S.O. Cyclorrhapha) Lonchopteridae  30
    Coleoptera (beetles) No further classification  21
    Scarabaeidae (chafers)  18
    Trichoptera (caddis flies) Limnephilidae 18
    Calliphoridae (blow flies)  6
    Coccinellidae (ladybirds) 6
    Diptera (S.O. Cyclorrhapha) Lauxaniidae 3
    _______________________________________________________________________________________________________________________
    E
    droppings collected at a sight in Tandragee 15km 9 miles from the shore of Lough Neagh
    Tipulidae (craneflies) .56%
    Nematocera eg midges and knats 1.12%
    Scathophagidae (dungflies) 11.86%
    Muscidae (house flies) 2.25%
    Cyclorrhapha eg circular seamed flies 7.90%
    Scarabaeidae (chafers) 23.44%
    Coccinellidae (ladybirds) .084%
    Coleoptera (beetles) No further classification 2.25%
    Cicadellidae (leafhoppers) 1.96%
    Hemiptera (true bugs) No further classification 3.95%
    Ichneumonidae (Ichneumons) 18.64%
    Cynapidae (gall wasps) 7.34%
    Pteromalidae eg parasitoiud wasps 5.64%
    Hymenoptera (wasps) No further classification 12.99%
    Comments from the person doing the analysis
    As at Birch Hill, five insect orders made up the diet of the swifts at Tandragee. However, the
    relative proportions were very different, and small wasps made up almost half the diet in this
    sample. Other important groups were cyclorrhapan flies, many of them dungflies, and beetles,
    almost all Serica sp. It appears that there were large hatchings of these small beetles in both
    areas during the summer of 2022. It is also noted that the Tandragee sample contained no
    aphid remains, thus confirming the finding at Birch Hill that aphid numbers were generally
    much lower in the area this year than last.
    ____________________________________________________________________________________________________________________
  • F
    Springhill, Moneymore 9km 6 miles from the shore of Lough Neagh
    Hemiptera (true bugs) No further classification 142
    Diptera (S.O.Nematocera mosquitoes, crane flies, gnats, black flies) No further classification 87
    Aphidae (aphids)  78
    Scathophagidae (dung-flies) 65
    Muscidae (house flies) 55
    Chironomidae (non-biting midges) 50
    Diptera (S.O. Cyclorrhapha) circular-seamed flies No further classification 50
    Diptera (S.O. Cyclorrhapha) Lonchopteridae  spear-winged flies 47
    Lepidoptera (moths) No further classification  43
    Coleoptera (beetles) No further classification  32
    Scarabaeidae (chafers) 20
    Tipulidae (craneflies) 6
    Cynapidae (gall wasps) 6
    Diptera (S.O. Cyclorrhapha) Phoridae hump-backed flies 4
    Psyllidae (jumping plant lice) 3
    _________________________________________________________________________________________________________________________
    What do house martins eat? 2022
    As with the swifts the droppings were collected below nests, dried and sent for analaysis

    Bibionidae

    St Mark’s flies

    3.85%

    Cyclorrhapha

    circular-seamed flies

    7.07%

    Cynapidae

    gall wasps

    34.4%

    Ichneumonidae

    ichneumon wasps

    9%

    Nematocera

    biting flies

    23.79%

    Scarabaeidae

    chafers

    17.36%

    Sciaridae

    dark-winged fungus gnats

    3.21%

    Tipulidae

    craneflies

    1.28%

    Comments from the person doing the analysis
    Compared to swifts (from a nearby location?), there was a higher percentage of small
    nematoceran flies in the diet of the house martins. This may have been because these small,
    relatively weak flying insects were more available at the lower height where house martins
    forage. There was also a high percentage of small wasps, similar to that in the swifts except
    in that the wasps eaten by the house martins were noticeably smaller. The majority of the
    fragments recovered were from Cynapidae (gall wasps) with body length <5mm. Again, this
    could have been because smaller insects were more available at lower altitude, or it may
    simply reflect the small body size of house martins. ________________________________________________________________________________________________________________________________
    Greenmount College/CAFRE, Antrim 2023

    Coleoptera (beetles) 

    chafers

    63%

    Hemiptera (true bugs) 

    No further classification 

    21%

    Diptera (S.O. Cyclorrhapha)

    No further classification

    13.3%

    Hemiptera (true bugs) 

    aphids

    11%

    Diptera (S.O. Cyclorrhapha) 

    dung-flies

    6.6%

    Trichoptera (caddis flies) 

    No further classification 

    6.33%

    Hymenoptera (wasps) 

    Ichneumons 

    5.3%

    Hemiptera (true bugs) 

    capsid bugs

    4.66%

    Hymenoptera (wasps) 

    gall wasps

    4%

    Diptera (S.O. Cyclorrhapha) 

    Dolichopodidae 

    3.3%

    Coleoptera (beetles) 

    No further classification 

    3.3%

    Diptera (S.O.Nematocera) 

    craneflies

    2.66%

    Neuroptera (lacewings) 

    brown lacewings

    2.66%

    Hymenoptera (wasps) 

    No further classification 

    2%

    Neuroptera (lacewings) 

    No further classification 

    2%

    Diptera (S.O. Cyclorrhapha) 

    house flies

    1.33%

    Lonchopteridae

     spear-winged flies

    1%

    Diptera (S.O.Nematocera) 

    Chironomidae

    1%

    Drosophilidae

    fruit flies

    .66%

     

     

    300 fragments

     Comments 

    The main insect groups in the sample were the same as those in the 2019 and 2021 reports although, again, the relative proportions varied. As last year, the two most important prey groups were Hemiptera (true bugs, including aphids) and cyclorraphan flies; the proportions of these two groups were broadly similar over the two years. Interestingly, the percentage of nematoceran flies has dropped again this year, and is now similar to that recorded in 2019 - this could well have been due to the long dry spell in early summer 2023. The number of beetles recorded was down this year. As discussed last year, most of the beetles were small chafers (Serica spp) which hatch in large numbers at intervals during the summer. Results suggest there were fewer hatches (or smaller ones) this year.  

    The most interesting findings were the presence of two new groups this year. Remains were found of caddis flies, which were almost certainly caught over water, since these insects have aquatic larvae. This suggests the birds were foraging at least some of the time over or close to the lough. The presence of remains of lacewings is unusual. These are not common insects, and they are delicately built and not strong fliers. I would not therefore have expected them to be present at the height at which swifts forage. It is likely they were carried on upcurrents during the hot, dry spell. __________________________________________________________________________________________________________________________________________________________

    Bolus from Caulside, Antrim 2023

    While out ringing swift chicks in July a bolus was found below the nest boxes. This gave us a perfect oportunity to see where the swift was in the previous 45-60 minutes

    Tipulidae (craneflies) 11.9%
    Ichneumonidae (Ichneumons) 9.7%
    Hemiptera (true bugs) No further classification 9.1%
    Hymenoptera (wasps) No further classification 8.6%
    Miridae (capsid bugs) 7%
    Scarabaeidae (chafers) 6.5%
    Diptera (S.O. Cyclorrhapha) No further classification 5.9%
    Chironomidae (chironomids) 5.4%
    Culicidae (mosquitos) 5.4%
    Aphidae (aphids) 5.4%
    Cynapidae (gall wasps) 4.8%
    Diptera (S.O.Nematocera) No further classification 4.3%
    Scathophagidae (dung-flies) 3.8%
    Muscidae (house flies) 3.8%
    Cecidomyiidae (gall midges) 3.2%
    Dolichopodidae (long-legged flies) 2%
    Drosophilidae (fruit flies) 2%
    Hymenoptera (wasps) Pteromalidae .5%

    Total number of identifiable fragments 185

    ___________________________________________________________________________________________________________

    Comments
    Essentially, the groups in the current analysis were similar to those found in chick faecal
    pellets locally in 2019 and 2022, although, as before, the relative proportions varied. This
    could have been due to the bolus being found earlier in the summer than were the pellets
    previously collected. Insect groups vary in how quickly numbers increase during the summer
    and also in how they react to prevailing weather conditions. However, it is also possible that
    differences could have arisen due to the different sampling methods used. Insect remains in
    the bolus were much less crushed and fragmented than were those in faecal pellets, and this
    would have meant that fragments of delicate, easily damaged insects such as nematoceran
    flies (craneflies and chironomids in this case) would survive better. This could explain why
    Nematocera formed a larger percentage of the current sample. The lower proportion of
    beetles (small chafers) is not unexpected. As explained previously, these beetles rise in large
    hatches periodically during the summer and so are only sporadically available in large
    numbers.
    It should also be noted that the bolus sample was smaller than the previous samples of 20
    pellets each. Therefore, the percentage frequency figures are likely to be slightly less
    accurate, although a sample containing 185 fragments is likely still to be useful in assessing
    the diet. Overall, it looks as if the swift which collected this bolus was feeding in much the
    same area and on roughly the same insect population as the birds in the previous studies.

  • __________________________________________________________________________________________________________________
    A side by side comparison. nfc or e.g. is no further classification
    Birch Hill  Birch Hill Birch Hill Birch Hill Birch Hill bolus '23 Birch Hill Crescent Arts  Dervock Springhill Kinawley Tandragee H martin CAFRE 
    2015 2019 2021 2022 2023 2024 2014 2018 2019 2019 2022 2022 2023
     
    Aphidae - greenfly/blackfly 18.20% 18% 17.54% 16.75% 10.50% 18.50% 10.90% 4.80% 11.33% 5.42% 11%
    Bibionidae e.g. St Mark’s flies 3.85%
    Cercopidae - froghoppers 5.29% 4.86%
    Chironomids - non biting midges  13.90% 25.70% 0.99% 0.13% 6% 18.50% 16.80% 2.63% 7.26% 5.74 1%
    Cicadellidae - leafhoppers 2.90% 3.64% 1.62% 2.94% 1.96%
    Coccinellidae - ladybirds 2.31% 1.08% 0.90% 0.95% 0.84%
    Coleoptera - water beetles 11.10%
    Coleoptera (beetles) nfc 1.65% 8.37% 0.75% 1.60% 6.51% 0.43% 3.34% 2.25% 3%
    Cynapidae - gall wasps 4.40% 2.81% 4.59% 6.73% 19.70% 0.90% 2.94% 0.87% 7.34% 34.40% 4%
    Cyclorrhapha eg circular-seamed flies 1.32% 9.72% 10.70% 0.60% 3.56% 7.26% 7.90% 7.07%
    Cyclorrhapha - nfc 1.60% 6% 19.29% 13.30%
    Chopodidae - long-legged flies 0.40% 3.30%
    Culicidae (mosquitoes) 0.74%
    Delphacidae - plant hoppers 0.90%
    Dolychopodidae - long-legged flies 0.50%
    Drosophila - fruit flies 0.66%
    Dytiscidae - water beetles 0.30%
    Gasteruptiidae - ?parasitic wasps 0.66%
    Hemerobiidae - brown lacewings 0.20% 3.10% 2.66%
    Hemerobiidae - lacewings nfc 2%
    Hemiptera (true bugs) nfc 0.80% 2.10% 16.50% 8.10% 20.20% 12.30% 12% 10.54% 6.10% 15.94% 3.95% 21%
    Hydroptilidae - water beetles 4.47% 6.75% 0.60% 12.99%
    Hymenoptera (wasps) nfc 4.47% 6.75% 9% 0.80% 1.24% 12.99% 2%
    Ichneumonidae - Ichneumons 7.80% 15.72% 5.67% 14.80% 7.60% 14.26% 18.64% 5.30%
    Lauxaniidae - acalyptrate flies 3.74% 0.47%
    Lepidoptera - moths 1% 0.75% 0.90% 1.39% 6.26%
    Limnephilidae - caddisflies 2.87%
    Miridae - capsid bugs 4.45% 4.66%
    Lonchopteridae - spear-winged flies 3.20% 11.5%% 2.15% 6.73% 6.80% 1.24% 6.83% 4.78% 1%
    Muscidae - house flies 6.45% 3.51% 2.50% 11.93% 7.99% 6.37% 2.25%
    Muscids / Calliphorids - stable / blow flies 6.20% 0.40% 1.35% 7.40% 2.01% 3.20% 1.33%
    Nematocera eg midges, gnats, mosquitoes 3.51% 7.40% 9.61% 12.64% 10.36% 1.12% 23.79%
    Odonata - damselflies 0.50%
    Opomyzidae - acalyptrate Diptera 0.77%
    Phoridae - hump-backed flies 0.80% 0.60% 0.58%
    Platygasterodae - parasitoid wasps 0.03% 4.60%
    Psyllidae - jumping plant lice 0.66% 7.30% 1.39% 0.43%
    Pteromalidae - parasitoid wasps 5.64%
    Sciaridae e.g. dark-winged fungus gnats 3.21%
    Scarabaeidae - chafers 8.70% 11.53% 10.54% 10.22% 3.30% 11.78% 2.87% 23.44% 17.36% 63%
    Scathophagidae - dung flies 10.20% 0.30% 4.80% 8.91% 3.49% 11.80% 9.44% 11.86% 6.60%
    Scatopsidae - dung midges 0.46%
    Sciaridae - fungus gnats 0.70%
    Sciomyzidae - marsh flies 3.74% 1.70% 0.77%
    Syrphidae - hoverflies 11.6%% 0.16% 0.46%
    Tipulidae - craneflies 0.10% 1.32% 4.05% 1.70% 0.87% 5.90% 0.56% 1.28% 2.66%
    Trichoptera - caddisflies 1.80% 5.58% 7.65% 6.33%